Oh my goodness! Unless you are a Tree of Life developer, you really shouldn't be here. This page is part of our beta test site, where we develop new features for the ToL, often messing up a thing or two in the process. Please visit the official version of this page, which is available here.
Under Construction

Lutrochidae

Lutrochus

Travertine beetles

William Shepard
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
Containing group: Byrrhoidea

Introduction

Lutrochidae is one of the smaller beetle families with only 11 species described in the one genus Lutrochus (Shepard 2002). However, Lutrochus will likely be split, and many other species and 2-3 additional genera await description. So far, all lutrochids occur only in the New World, from the USA south at least to Paraguay (Shepard and Julio 2010).

All lutrochids are stream-associated.  One group of species is often found in bankside vegetation hanging into or over the water or in leaf packs that accumulate on sticks and rocks. A second group of species occurs where artesian waters enter streams and travertine deposits form on submerged wood and vegetation. Larvae of this group mine the travertine consuming the organic matter. The common name for the family comes from this group of species.

Known life cycles are univoltine. Oviposition is into wet organic matter. Larvae undergo several stadia and then pupate under objects just above the water level. Adults are found close to the larvae. Adults and larvae feed on algae and waterlogged organic matter. Life cycle information is available for L. arizonicus (Brown and Murvosh 1970) and L. germari (Costa et al. 1996).

Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window

Lutrochus. © Alex Wild

Characteristics

Adult and larval morphologies have been well described (Hinton 1939; Brown and Murvosh 1970; Brown 1991; Costa et al. 1996).  Identification keys exist for North American species (Brown 1972) and most South American species (Grouvelle 1896) (in French).

Adults can be recognized by the following characters:

Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window

Lutrochus vestitus, larva © Susan McCormick

Larvae can be recognized by the following characters:

Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window

Lutrochus luteus, larva © Susan McCormick

Classification

Lutrochids were first placed in Dryopidae. Then Hinton (1939) moved them into Limnichidae where they remained until given their own family status (Kasap and Crowson 1975).

 Identification keys exist for North American species (Brown 1972) and most South American species (Grouvelle 1896) (in French).

Relationships of Lutrochidae to Other Beetles

Phylogenetic analyses of morphological data have put Lutrochidae as a sister group to Dryopidae (Lawrence 1987), to (Elmidae + Psephenidae) (Beutel 1995) or to (Limnichidae + Dryopidae + Heteroceridae) (Costa et al. 1999).

Other Names for Lutrochus

References

Beutel, R. G. 1995. Phylogenetic analysis of Elateriformia (Coleoptera: Polyphaga) based on larval characters. Journal of Zoological Systematics and Evolutionary Research, 33(4): 145-171.

Brown, H. P. 1972. Aquatic Dryopoid Beetles (Coleoptera) of the United States. U. S. Environmental Protection Agency. Biota of Freshwater Ecosystems. Identification Manual No. 6. 82 pp.

Brown, H. P. 1991. Lutrochidae (Dryopoidea). Pages 397-399. In: F. W. Stehr (ed.). Immature Insects. Volume 2. Kendall/Hunt. Dubuque, Iowa. 975 pp.

Brown, H. P. and C. Murvosh. 1970. Lutrochus arizonicus new species, with notes on ecology and behavior (Coleoptera, Dryopoidea, Limnichidae). Annals of the Entomological Society of America, 63(4): 1030-1035.

Costa, C., I. Serfio, S. A. Vanin and É. P. Teixeira. 1996. Larvae of Neotropical Coleoptera. XXIII: Lutrochus germari Grouvelle, description of immatures, redescription of adult and bionomics (Dryopoidea, Lutrochidae). Revista Brasileira de Entomologia, 40(1): 47-56.

Costa, C., S. A. Vanin and S. Ide. 1999. Systematics and bionomics of Cneoglossidae with a cladistic analysis of Byrrhoidea sensu Lawrence & Newton (1995) (Coleoptera, Elateriformia). Arqivos de Zoologia, 35(3): 231-300.

Grouvelle, A. 1896. Description de deux Dryopides du genre Lutrochus Er. Avec tableau general des espèces [Col.]. Bulletin de la Société de France, 1-2: 16-17.

Hinton, H. E. 1939. An inquiry into the natural classification of the Dryopoidea, based partly on a study of their internal anatomy (Col.). Transactions of the Royal Entomological Society of London, 89(7): 133-184.

Ide, S., C. Costa and S. A. Vanin. 2005. 18.4. Lutrochidae Kasap & Crowson, 1975. Pages 508-512. In: R. G. Beutel and R. A. B. Leschen (eds.). Handbook of Zoology. Volume IV. Arthropoda: Insecta. Coleoptera, Beetles. Volume 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). Walter de Gruyter. Berlin. 567 pp.

Lawrence, J. F. 1987. Rhinorhipidae, a new beetle family from Australia, with comments on the phylogeny of the Elateriformia. Invertebrate Taxonomy, 2: 1-53.

Shepard, W. D. 2002. Chapter 45. LUTROCHIDAE Kasap and Crowson 1975. Pages 123-124. In: Arnett, R. H., M. C. Thomas, P. E. Skelley and J. H. Frank (eds.). American Beetles. Volume 2. CRC Press. Boca Raton, Florida. 861 pp.

Shepard, W. D. and C. A. Julio. 2010. Estudio preliminary de las familias de escarabajos acuáticos Dryopidae, Elmidae, Lutrochidae y Psephenidae conocidos de Paraguay (Coleoptera: Byrrhoidea). Boletin del Museo Nacional de Historia Natural de Paraguay, 16(1): 30-42.

Information on the Internet

Title Illustrations
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
Scientific Name Lutrochus
Location USA AZ Yavapai County: Red Rock Crossing, 34º49'N 111º48'W, 1200m
Source Lutrochus sp. - Travertine Beetles
Copyright © Alex Wild
About This Page


Univerrsity of California, Berkeley

Correspondence regarding this page should be directed to William Shepard at

Page: Tree of Life Lutrochidae. Lutrochus. Travertine beetles. Authored by William Shepard. The TEXT of this page is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Shepard, William. 2011. Lutrochidae. Lutrochus. Travertine beetles. Version 26 February 2011 (under construction). http://tolweb.org/Lutrochus/9117/2011.02.26 in The Tree of Life Web Project, http://tolweb.org/

edit this page
close box

This page is a Tree of Life Branch Page.

Each ToL branch page provides a synopsis of the characteristics of a group of organisms representing a branch of the Tree of Life. The major distinction between a branch and a leaf of the Tree of Life is that each branch can be further subdivided into descendent branches, that is, subgroups representing distinct genetic lineages.

For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page.

close box

Lutrochus

Page Content

articles & notes

collections

people

Explore Other Groups

random page

  go to the Tree of Life home page
top