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Hesperiidae Latreille 1809


Andrew V. Z. Brower and Andrew Warren
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  Checkered skipper butterfly
taxon links [up-->]Trapezitinae [up-->]Eudaminae [up-->]Hesperiinae [up-->]Coeliadinae [up-->]Euschemoninae [up-->]Pyrginae [up-->]Heteropterinae [down<--]Ditrysia Interpreting the tree
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The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

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Containing group: Ditrysia

Discussion of Phylogenetic Relationships

The hypothesis of relationships is based on the combined morphological/molecular cladistic analysis of Warren et al. (2008). Eudaminae is removed from Pyrginae in order to preserve monophyly of the latter. Pyrrhopygini (formerly viewed as a subfamily) is nested within Pyrginae.

Other Names for Hesperiidae Latreille 1809


Ackery, P. R., R. de Jong, and R. I. Vane-Wright. 1999. The butterflies: Hedyloidea, Hesperioidea, and Papilionoidea. Pages 264-300 in: Lepidoptera: Moths and Butterflies. 1. Evolution, Systematics, and Biogeography. Handbook of Zoology Vol. IV, Part 35. N. P. Kristensen, ed. De Gruyter, Berlin and New York.

Warren, A. D., Ogawa, J. R. & Brower, A. V. Z. 2008. Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperiodea). Cladistcs 24, 642-676.

Information on the Internet

Title Illustrations
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Scientific Name Hylephila phyleus
Specimen Condition Live Specimen
Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 2.5.
Copyright © Bruce Marlin
Scientific Name Atarnes sallei
Location El Naranjo, San Luis Potosi, Mexico
Specimen Condition Live Specimen
Source Orange-spotted Skipper
Source Collection Neotropical Butterflies
Copyright © 2002 Will and Gill Carter
Checkered skipper butterfly
Scientific Name Pyrgus communis
Location Mount Bruno, elevation 1300 feet (San Mateo County, California, USA)
Comments Checkered skipper butterfly
Creator Photograph by T. W. Davies
Specimen Condition Live Specimen
Source Collection CalPhotos
Copyright © 1999 California Academy of Sciences
About This Page

Middle Tennessee State University, Murfreesboro, Tennessee, USA

University of Florida, Gainesville, Florida, USA

Correspondence regarding this page should be directed to Andrew V. Z. Brower at and Andrew Warren at

Page: Tree of Life Hesperiidae Latreille 1809. Skippers. Authored by Andrew V. Z. Brower and Andrew Warren. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Brower, Andrew V. Z. and Andrew Warren. 2008. Hesperiidae Latreille 1809. Skippers. Version 07 April 2008 (under construction). http://tolweb.org/Hesperiidae/12028/2008.04.07 in The Tree of Life Web Project, http://tolweb.org/

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For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page.

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