Oh my goodness! Unless you are a Tree of Life developer, you really shouldn't be here. This page is part of our beta test site, where we develop new features for the ToL, often messing up a thing or two in the process. Please visit the official version of this page, which is available here.
Under Construction

Euptychiina Reuter 1896

Andrew V. Z. Brower
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
taxon links [up-->]Coeruleotaygetis [up-->]Pindis squamistriga [up-->]Cercyeuptychia [up-->]Megisto [up-->]Forsterinaria [up-->]Prenda clarissa [up-->]Erichthodes [up-->]Taygetina [up-->]Neonympha [up-->]Splendeuptychia [up-->]Caenoptychia boulleti [up-->]Caeruleuptychia [up-->]Hermeuptychia [up-->]Godartiana [up-->]Palaeonympha opalina [up-->]Amphidecta [up-->]Taydebis [up-->]Praefaunula [up-->]Capronnieria [up-->]Moneuptychia [up-->]Zischkaia [up-->]Megeuptychia [up-->]Taygetis [up-->]Pseudeuptychia [up-->]Cissia [up-->]Posttaygetis [up-->]Paryphthimoides [up-->]Parataygetis [up-->]Carminda [up-->]Taygetomorpha [up-->]Chloreuptychia [up-->]Euptychia [up-->]Paramacera [up-->]Guaianaza [up-->]Yphthimoides [up-->]Pareuptychia [up-->]Archeuptychia [up-->]Satyrotaygetis [up-->]Euptychiina incertae sedis [up-->]Cyllopsis [up-->]Harjesia [up-->]Rareuptychia [up-->]Euptychoides [up-->]Cepheuptychia [up-->]Pseudodebis [up-->]Pharneuptychia [up-->]Magneuptychia Not MonophyleticNot MonophyleticNot MonophyleticMonophyly UncertainMonophyly UncertainMonophyly UncertainPhylogenetic position of group is uncertain and group is not monophyleticPhylogenetic position of group is uncertain and group is not monophyletic[down<--]Satyrini Interpreting the tree
close box

This tree diagram shows the relationships between several groups of organisms.

The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

example of a tree diagram

You can click on the root to travel down the Tree of Life all the way to the root of all Life, and you can click on the names of descendent subgroups to travel up the Tree of Life all the way to individual species.

For more information on ToL tree formatting, please see Interpreting the Tree or Classification. To learn more about phylogenetic trees, please visit our Phylogenetic Biology pages.

close box
Containing group: Satyrini


With the exception of the Taiwanese Palaeonympha, Euptychiina are an exclusively New World, mainly neotropical satyrine clade. A few species and genera extend into the U. S., including Megisto, Hermeuptychia and Cyllopsis, but the vast preponderance of species occur in tropical Central and South America. Most species are monocot feeders, but a few, such as members of Euptychia, feed on unusual hosts such as the spikemoss Selaginella.

Discussion of Phylogenetic Relationships

It is becoming increasingly apparent (e. g., Murray and Prowell 2005; Peña et al. 2010) that many euptychiine genera are para- or polyphyletic, and significant effort will need to be invested to sort out relationships among members of this clade. Some putative sister taxa are identified in the above tree based mainly on Peña et al. (2006, 2010), but even these hypotheses are in need of corroboration.


Forster W. 1964. Beiträge zur Kenntnis der Insektenfauna Boliviens. XIX. Einleitung Lepidoptera III. Satyridae. Veröffentlichungen der zoologischen Staatsammlung München 3: 81-160, plates 25-33.

Lamas G ed. 2004. Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea - Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.

Miller LD. 1968. The higher classification, phylogeny and zoogeography of the Satyridae (Lepidoptera). Mem. Amer. Entomol. Soc. 24: 1-174.

Murray D, and Prowell DP. 2005. Molecular phylogenetics and evolutionary history of the neotropical satyrine subtribe Euptychiina (Nymphalidae: Satyrinae). Mol. Phylogenet. Evol. 34: 67-80.

Peña C, Nylin S, Freitas AVL and Wahlberg N. 2010. Biogeographical history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta 39: 243-258.

Peña C, Wahlberg N, Weingartner E, Kodandaramaiah U, Nylin S, Freitas AVL, and Brower AVZ. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Mol. Phylogenet. Evol. 40: 29-49.

Title Illustrations
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
Scientific Name Cyllopsis gemma
Location Chattahoochee Nat'l Rec Area, Medlock Bridge, Georgia, USA
Specimen Condition Live Specimen
Source Gemmed Satyr - Cyllopsis gemma
Source Collection BugGuide.Net
Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 2.0.
Copyright © 2002 Troy Bartlett
Scientific Name Megisto cymela
Location Arlington, Texas, USA
Specimen Condition Live Specimen
Source Butterfly away?
Source Collection Flickr
Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial-NoDerivs License - Version 2.0.
Copyright © 2006 DeWaun Simmons
Scientific Name Cepheuptychia cephus
Location Manu, Peru
Specimen Condition Live Specimen
Identified By Kim Garwood
Sex Female
Life Cycle Stage adult
View ventral
Source Cephus Satyr
Source Collection Neotropical Butterflies
Copyright © 2004 Kim Garwood
About This Page

Middle Tennessee State University, Murfreesboro, Tennessee, USA

Correspondence regarding this page should be directed to Andrew V. Z. Brower at

Page: Tree of Life Euptychiina Reuter 1896. Authored by Andrew V. Z. Brower. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Brower, Andrew V. Z. 2011. Euptychiina Reuter 1896. Version 16 May 2011 (under construction). http://tolweb.org/Euptychiina/70791/2011.05.16 in The Tree of Life Web Project, http://tolweb.org/

edit this page
close box

This page is a Tree of Life Branch Page.

Each ToL branch page provides a synopsis of the characteristics of a group of organisms representing a branch of the Tree of Life. The major distinction between a branch and a leaf of the Tree of Life is that each branch can be further subdivided into descendent branches, that is, subgroups representing distinct genetic lineages.

For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page.

close box


Page Content

articles & notes



Explore Other Groups

random page

  go to the Tree of Life home page