Biina
Bia
Andrew V. Z. Brower- Bia actorion
- Bia peruana
Discussion of Phylogenetic Relationships
Bia was an enigmatic genus for a long time. Miller (1968) placed it as a basal satyrine in Biinae, a subfamily including Antirrhea and Melanitis that is now viewed as polyphyletic. Brower (2000) showed that Bia belonged to Brassolini based on sequences of the wingless gene, and this has subsequently been supported by additional DNA sequence data (Peña et al. 2005) as well as morphology (Freitas et al. 2002; Vane-Wright and Boppré 2004).
References
Brower AVZ. 2000. Phylogenetic relationships among the Nymphalidae (Lepidoptera) inferred from partial sequences of the wingless gene. Proc. R. Soc. Lond. B 267: 1201-1211.
Casagrande, MM. 2004. Brassolini. In Lamas G ed. 2004. Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea - Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.
Freitas AVL, Murray D, and Brown KS, Jr. 2002. Immatures, natural history and the systematic position of Bia actorion (Nymphalidae). J. Lepid. Soc. 56: 117-122.
Miller LD. 1968. The higher classification, phylogeny and zoogeography of the Satyridae (Lepidoptera). Mem. Amer. Entomol. Soc. 24: 1-174.
Peña C, Wahlberg N, Weingartner E, Kodandaramaiah U, Nylin S, Freitas AVL, and Brower AVZ. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Mol. Phylogenet. Evol. 40: 29-49.
Vane-Wright RI, and Boppre M. 2004. Adult morphology and the higher classification of Bia Huebner (Lepidoptera: Nymphalidae). Bonner zoologische Beitrage 53: 235-254.
Title Illustrations

| Location | Rio Cristalino Lodge, Matto Grosso, Brasil | 
|---|---|
| Specimen Condition | Live Specimen | 
| Identified By | Gil Carter | 
| Life Cycle Stage | adult | 
| View | ventral | 
| Source | Actorion Owlet | 
| Source Collection | Neotropical Butterflies | 
| Copyright | © 2004 Gil Carter | 
About This Page
	
	
        
       
    
    
	 
 
    
	Middle Tennessee State University, Murfreesboro, Tennessee, USA
				Correspondence regarding this page should be directed to Andrew V. Z. Brower at 
            
Page copyright © 2010
    
	     Page: Tree of Life 
     Biina . Bia .  
    Authored by 
    Andrew V. Z. Brower.         
	The TEXT of this page is licensed under the 
    
    Creative Commons Attribution License - Version 3.0. Note that images and other media 
    featured on this page are each governed by their own license, and they may or may not be available 
    for reuse. Click on an image or a media link to access the media data window, which provides the 
    relevant licensing information. For the general terms and conditions of ToL material reuse and 
    redistribution, please see the Tree of Life Copyright 
    Policies.
    Page: Tree of Life 
     Biina . Bia .  
    Authored by 
    Andrew V. Z. Brower.         
	The TEXT of this page is licensed under the 
    
    Creative Commons Attribution License - Version 3.0. Note that images and other media 
    featured on this page are each governed by their own license, and they may or may not be available 
    for reuse. Click on an image or a media link to access the media data window, which provides the 
    relevant licensing information. For the general terms and conditions of ToL material reuse and 
    redistribution, please see the Tree of Life Copyright 
    Policies.
- First online 29 October 2006
- Content changed 13 February 2010
Citing this page:
Brower, Andrew V. Z. 2010. Biina . Bia . Version 13 February 2010 (under construction). http://tolweb.org/Bia/70748/2010.02.13 in The Tree of Life Web Project, http://tolweb.org/

 
			
	    
		 
        	
	
		 
			
	
		

 
     
 
	
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