Oh my goodness! Unless you are a Tree of Life developer, you really shouldn't be here. This page is part of our beta test site, where we develop new features for the ToL, often messing up a thing or two in the process. Please visit the official version of this page, which is available here.
Under Construction


Brian M. Wiegmann and David K. Yeates
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
Asilidae, robber fly
taxon links [up-->]Pantophthalmidae [up-->]Mydidae [up-->]Stratiomyidae [up-->]Therevidae [up-->]Evocoidae [up-->]Apioceridae [up-->]Apystomyiidae [up-->]Acroceridae [up-->]Oreoleptidae [up-->]Empidoidea [up-->]Athericidae [up-->]Apsilocephalidae [up-->]Xylomyidae [up-->]Xylophagidae [up-->]Bombyliidae [up-->]Hilarimorphidae [up-->]Cyclorrhapha Not MonophyleticMonophyly UncertainMonophyly UncertainPhylogenetic position of group is uncertain[down<--]Diptera Interpreting the tree
close box

This tree diagram shows the relationships between several groups of organisms.

The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

example of a tree diagram

You can click on the root to travel down the Tree of Life all the way to the root of all Life, and you can click on the names of descendent subgroups to travel up the Tree of Life all the way to individual species.

For more information on ToL tree formatting, please see Interpreting the Tree or Classification. To learn more about phylogenetic trees, please visit our Phylogenetic Biology pages.

close box
Tree adapted from Nagatomi and Liu 1994; Yeates 1994; Yeates 2002; Wiegmann et al. 2003; Yeates, Irwin, and Wiegmann 2003; Yeates, Meier, and Wiegmann 2003; Moulton and Wiegmann 2004; Zloty et al. 2005.
Containing group: Diptera


The suborder Brachycera represents a major division of the Diptera containing approximately 120 families and a great diversity of species, morphological innovations, and life history strategies. The name "Brachycera" or "shortened horn" refers to their shortened antennae—a reduced antenna with fewer than 8 antennal flagellomeres is the easiest to recognise of a list of defining features (see below). The origins of the Brachycera can be traced to the late Triassic or earliest Jurassic (245-208 mya) based on fossil evidence (Evenhuis 1995; Grimaldi and Cumming 1999; Yeates and Wiegmann 1999) and comparisons of evolutionary rates in DNA sequences (Wiegmann et al. 2003).


Brachycera is well supported monophyletic lineage supported by a number of morphological synapomorphies (Yeates 2002; Yeates and Wiegmann 1999; Sinclair et al. 1993; Woodley 1989):

Brachyceran flies exhibit a broad array of feeding strategies, life histories and behavioral patterns. Many of the orthorrhaphous lineages are scavengers, predators or parasitoids as larvae. Multiple major radiations of species diversity, feeding habits and habitat specialization can be found in the Cyclorrhapha or "higher" flies.

Discussion of Phylogenetic Relationships

Traditional non-phylogenetic classifications of the brachycera break the group into two major sections, the 'Orthorrhapha' and Cyclorrhapha. The 'Orthorrhaphous' Brachycera is a paraphyletic assemblage of all those families not included in the Cyclorrhapha. Recent classifications based on explicit phylogenetic evidence include 4 monophyletic infraorders: Stratiomyomorpha, Xylophagomorpha, Tabanomorpha, and Muscomorpha.

Brachyceran phylogeny has been the subject of intensive study in recent years. Comprehensive phylogenetic treatments in Hennig (1973), Griffiths (1972), Woodley (1989) and Sinclair et al. (1994) began the explicit definition of clades based on synapomorphic character states (see Yeates and Wiegmann 1999 for a review). Quantitative phylogenetic analyses of morphological (Yeates 1994, 2002; Yeates et al. 2003; Meier and Hilger 2000) and gene sequence data (Wiegmann et al. 2000, Wiegmann et al. 2003; Collins and Wiegmann 2001ab; Moulton and Wiegmann 2004) continue to add resolution and support to many of the key lineages of brachyceran flies.


Collins, K. P. and B. M. Wiegmann. 2002. Phylogenetic relationships and placement of the Empidoidea (Diptera: Brachycera) based on 28S rDNA and EF-1a sequences. Insect Syst. Evol. 33: 21-444.

Griffiths, G. C. D. 1972. The phylogenetic classification of Diptera Cyclorrhapha, with special reference to the male postabdomen. Series entomologica 8, 340pp. The Hague.

Griffiths, G. C. D. 1994. Relationships among the major subgroups of Brachycera (Diptera): A critical review. Can. Entomol. 126:861-880.

Hennig, W. 1973. Diptera. In: W. Kukenthal (ed.) Handbuch der Zoologie, IV: Arthropoda. de Gruyter, New York, pp. 1-337.

Krivosheina, N. P. 1991. Phylogeny of lower Brachycera (Diptera): A new view. Acta Entomol. Bohemoslov. 88:81-92.

McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth, and D. M. Wood (eds.). 1981, 1987. Manual of Nearctic Diptera, Vol. 1 & 2. Research Branch, Agriculture Canada, Monographs 27 & 28.

McAlpine, J. F. and D. M. Wood (eds.). Manual of Nearctic Diptera, Vol. 3. Research Branch, Agriculture Canada, Monograph 32.

Moulton, J. K. and B. M. Wiegmann. 2004. Evolution and phylogenetic utility of CAD (rudimentary) among Mesozoic-aged Eremoneuran Diptera (Insecta). Mol. Phylogenet. Evol. 31: 363-378.

Nagatomi, A. 1991. History of some families of Diptera, chiefly those of the lower Brachycera (Insecta: Diptera). Bull. Biog. Soc. Japan 46(1-22): 21-38.

Nagatomi, A. and N. Liu. 1994. Apystomyiidae, a new family of Asiloidea (Diptera). Acta Zool. Hung. 40:203-218.

Sinclair, B. J. 1992. A phylogenetic interpretation of the Brachycera (Diptera) based on the larval mandible and associated mouthpart structures. Syst. Entomol. 17:233-252.

Sinclair, B. J., Cumming, J. M. and D. M. Wood. 1994. Homology and phylogenetic implications of the male genitalia in Diptera-Lower Brachycera. Entomol. Scand. 24:407-432.

Wada, S. 1991. Morphological evidence for the direct sister group relationship between the Schizophora and the Syrphoidea (Aschiza) in the phylogenetic systematics of the Cyclorrhapha (Diptera: Brachycera). J. Nat. Hist. 25:1531-1570.

Wiegmann, B. M., C. Mitter, and F. C. Thompson. 1993. Evolutionary origin of the Cyclorrhapha (Diptera): tests of alternative morphological hypotheses. Cladistics, 9:41-81.

Wiegmann, B. M., D. K. Yeates, J. L. Thorne, and H. Kishino. 2003. Time flies, a new molecular time-scale for brachyceran fly evolution without a clock. Syst. Biol. 52:745-756.

Woodley, N. E. 1989. Phylogeny and classification of the "Orthorrhaphous" Brachycera. In: McAlpine J.F., Wood D.M. (eds.)Manual of Nearctic Diptera 3. Research Branch, Agriculture Canada, Monograph 32:1371-1395.

Yeates, D. K. 1994. The cladistics and classification of the Bombyliidae (Diptera: Asiloidea). Bull. Am. Mus. Nat. Hist. 219:1-191.

Yeates, D. K. 2002. Relationships of the lower Brachycera (Diptera): A quantitative synthesis of morphological characters. Zool. Scripta 31:105-121.

Yeates, D. K. and B. M. Wiegmann. 1999. Congruence and controversy: Toward a higher-level phylogeny of Diptera. Ann. Rev. Entomol. 44:397-428.

Yeates, D. K., M. E. Irwin, and B. M. Wiegmann. 2003. Ocoidae, a new family of asiloid flies (Diptera: Brachycera: Asiloidea), based on Ocoa chilensis gen. and sp. nov. from Chile, South America. Syst. Entomol. 28:417-431.

Yeates, D. K., R. Meier, and B. M. Wiegmann. 2003. Phylogeny of true flies (Diptera): A 250 million year old success story in terrestrial diversification. Entomol. Abh. 61:119.

Zaitsev, V. F. 1991. On the phylogeny and systematics of the dipteran superfamily Bombylioidea (Diptera). Entomologicheskoe Obozrenie 70(3): 716-736.

Zloty, J., B. J. Sinclair, & G. Pritchard. 2005. Discovered in our backyard: a new genus and species of a new family from the Rocky Mountains of North America (Diptera, Tabanomorpha). Systematic Entomology. 30:248-266.

Title Illustrations
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
Asilidae, robber fly
Scientific Name Asilidae
Location New Mexico, USA
Specimen Condition Live Specimen
Identified By Alex Wild
Life Cycle Stage Adult
Copyright © 2005 Alex Wild
Scientific Name Lamprogaster sp.
Comments An Australian platystomatid fly, (Diptera: Brachycera: Cyclorrhapha: Platystomatidae)
Specimen Condition Dead Specimen
Identified By D. K. Yeates
Copyright © 2004 David McClenaghan
About This Page

Brian M. Wiegmann
North Carolina State University, Raleigh, North Carolina, USA

CSIRO Division of Entomology, Canberra, Australia

Correspondence regarding this page should be directed to Brian M. Wiegmann at and David K. Yeates at

All Rights Reserved.

Citing this page:

Wiegmann, Brian M. and David K. Yeates. 2007. Brachycera. Version 29 November 2007 (under construction). http://tolweb.org/Brachycera/10500/2007.11.29 in The Tree of Life Web Project, http://tolweb.org/

edit this page
close box

This page is a Tree of Life Branch Page.

Each ToL branch page provides a synopsis of the characteristics of a group of organisms representing a branch of the Tree of Life. The major distinction between a branch and a leaf of the Tree of Life is that each branch can be further subdivided into descendent branches, that is, subgroups representing distinct genetic lineages.

For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page.

close box


Page Content

articles & notes




Explore Other Groups

random page

  go to the Tree of Life home page